Alexander Grayson becomes drawn to a woman who looks like his dead former love, while her fiance, Harker, worries about his ability to give her the life she deserves.
Grayson and Lady Jayne become lovers as he looks into her connections; Grayson helps Mina face a challenge; Jonathan strikes a bargain that has unexpected consequences.
Grayson investigates whether Lady Jayne is a vampire hunter; Lucy employs absinthe, romance and a tour of London to mend Nina's broken heart; Grayson earns a powerful enemy.
Grayson tries to win Lady Jayne's heart; Lucy hides her broken heart behind planning Mina and Harker's engagement party; Grayson's reunion with an old friend complicates matters; Van Helsing suffers a setback.
Harker learns Gen. Shaw isn't the hero he pretends to be; Van Helsing's first test with the solar serum is a failure; Davenport's people abduct Renfield; when Grayson dances with Mina at her engagement party, their feelings are revealed.
Browning hosts a board meeting in the sunlight in an effort to expose Grayson; Grayson and Van Helsing acquire a vital component for the wireless electricity technology; Mina and Harker advance their relationship.
Grayson is intoxicated by his invulnerability to sunlight; Renfield is sent to Budapest; Harker realizes he was manipulated.
The cancellation of Grayson and Van Helsing's demonstration leads Van Helsing to seek revenge; Harker forbids Mina from seeing Grayson; Lord Davenport's men attack Mina; Lady Jane learns Dracula is in London.
While Mina recovers, Harker decides to join the Order of the Dragon; Browning searches for his children; Lady Jane gets ready for the ultimate vampire hunt.
A public demonstration of Grayson's technology has terrible consequences; Lucy joins the undead; Van Helsing's revenge against Mr. Browning and Lady Jane leads to a confrontation with Grayson; Mina discovers the secret of Grayson's love.
In a Camelot flashback, Hook becomes a Dark One and his lust for revenge against Rumplestiltskin is reignited. When Emma and Hook disagree about their best course of action, the final pieces of the puzzle are revealed as events in Camelot catch up to the present and we witness a charged confrontation between the forces of light and dark that sends our heroes on a collision course with destiny. Meanwhile, in Storybrooke, Hook’s centuries old lust for revenge against Gold puts both of their fates